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It's the wing's call on the side that the ball goes to. His primary responsibility is that wideout. If he doesn't break downfield, he'll stay put. The other wing is downfield on a pass if his guy goes. If there is no one split out to his side, he'll drift there anyway after he reads his tackle key.
We have a mechanic which half the R's want and half don't where we 'punch' backward on a backward pass, the same as we would on an eligible off the line closest to us before the snap. The other half just lets the play go, no whistle and waits for everyone to catch on if there is a loose ball (the wideout drops it).
If the wideout takes off and the wing goes with him and, say, a slot guy steps back, the R has the call. The rule of thumb is that if its close, its an incomplete pass.
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