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Yes, the defense can appeal. The defense's err was during continous action of the play, throwing the ball out of play, not a post action appeal. Once the ball goes out of play, R1 is allowed to complete his base running responsibilities. That is he should have retouched 1B, then continued onto 2 and 3b. He left the appeal open by not fulfilling his baserunning responsibilities.
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Its' not a matter of being right or wrong, it's a matter of working hard to get it right.
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