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Only the first lane spaces are restricted to defense only. The last two spaces are for the defense if they want them, and the middle two are for the offense if they want them. If the preferred occupants decline to occupy those four spaces, the other team may take them if they desire.
Note the difference in wording between article 4c and the next two, 4d and 4e. 4c uses "shall" and the other two use "may."
Not only may the offense not use the the first spaces if the defense declines, the defense may not decline to use them.
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